:root {
  --ink: #1f241d;
  --muted: #626b5e;
  --line: #dfd8c9;
  --paper: #f3f4f2;
  --panel: #fffdf8;
  --cream: #f8f6f1;
  --accent: #b48a3c;
  --accent-2: #2f5e3b;
  --accent-3: #6c9b53;
  --soft: #eef1e9;
  --warn: #f4ead2;
  --radius: 12px;
  --shadow: 0 10px 28px rgba(41, 45, 37, 0.075);
  --hero-image: url("assets/images/habitat-libelloides.jpg");
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  background:
    radial-gradient(circle at 20% 0, rgba(180, 138, 60, 0.09), transparent 36vw),
    linear-gradient(180deg, var(--cream), var(--paper));
  color: var(--ink);
  font-family: ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  line-height: 1.5;
}

a {
  color: inherit;
}

.site-hero {
  position: relative;
  min-height: 164px;
  overflow: hidden;
  background:
    linear-gradient(90deg, rgba(14, 24, 15, 0.86), rgba(26, 45, 25, 0.48), rgba(21, 29, 18, 0.28)),
    var(--hero-image) center / cover;
  color: #fffdf8;
}

.hero-overlay {
  position: absolute;
  inset: 0;
  backdrop-filter: blur(1.4px);
  background:
    linear-gradient(180deg, rgba(7, 16, 8, 0.16), rgba(7, 16, 8, 0.06)),
    rgba(20, 28, 20, 0.12);
}

button {
  min-height: 38px;
  border: 1px solid var(--line);
  border-radius: 12px;
  background: var(--panel);
  color: var(--ink);
  padding: 8px 14px;
  font: inherit;
  text-decoration: none;
  cursor: pointer;
}

button:hover,
button:focus-visible {
  border-color: var(--accent-2);
  outline: 0;
}

.hero-content {
  position: relative;
  z-index: 1;
  display: flex;
  align-items: flex-start;
  justify-content: space-between;
  gap: 24px;
  width: min(1480px, calc(100% - 48px));
  margin: 0 auto;
  padding: clamp(18px, 3.6vw, 28px) 0;
}

.hero-content h1 {
  margin: 0;
  max-width: 980px;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(2.1rem, 4.4vw, 3.8rem);
  line-height: 0.98;
  letter-spacing: 0;
  text-shadow: 0 8px 24px rgba(0, 0, 0, 0.34);
}

.hero-content p:not(.eyebrow) {
  max-width: 780px;
  margin: 12px 0 0;
  font-size: clamp(1rem, 1.8vw, 1.2rem);
  color: rgba(255, 253, 248, 0.92);
}

.hero-content strong {
  display: inline-block;
  margin-top: 8px;
  color: #cbe889;
  font-size: 1.05rem;
}

.hero-brand {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: 6px;
  margin-top: 3px;
  color: #fffdf8;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(0.82rem, 1vw, 1rem);
  font-weight: 800;
  line-height: 1.05;
  letter-spacing: 0.08em;
  text-decoration: none;
  text-transform: uppercase;
  white-space: nowrap;
  text-align: center;
}

.hero-brand img {
  display: block;
  width: clamp(70px, 7.5vw, 102px);
  height: auto;
  max-height: none;
  filter: drop-shadow(0 6px 18px rgba(0, 0, 0, 0.32));
}

.brand-mark {
  display: grid;
  place-items: center;
  width: 58px;
  height: 58px;
  border: 2px solid rgba(255, 255, 255, 0.72);
  border-radius: 16px;
  background: rgba(255, 255, 255, 0.13);
  font-family: ui-sans-serif, system-ui, sans-serif;
  font-size: 1rem;
  letter-spacing: 0;
}

.step-nav {
  position: sticky;
  top: 0;
  z-index: 20;
  display: grid;
  grid-template-columns: auto auto auto auto 1fr auto;
  align-items: center;
  gap: 10px;
  padding: 8px clamp(10px, 2vw, 20px);
  border-bottom: 1px solid rgba(209, 199, 180, 0.72);
  background: rgba(248, 246, 241, 0.92);
  backdrop-filter: blur(14px);
}

.step-pill {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 9px;
  min-height: 40px;
  border: 1px solid var(--line);
  border-radius: 999px;
  background: rgba(255, 253, 248, 0.86);
  color: var(--ink);
  padding: 7px 13px;
  font-weight: 800;
  text-decoration: none;
  box-shadow: 0 8px 22px rgba(40, 48, 38, 0.05);
}

.step-pill.active {
  border-color: transparent;
  background: var(--accent-2);
  color: #fff;
}

.step-number,
.nav-icon {
  display: grid;
  place-items: center;
  width: 24px;
  height: 24px;
  border-radius: 50%;
  background: rgba(47, 94, 59, 0.12);
  color: currentColor;
  font-size: 0.82rem;
}

.active .step-number {
  background: rgba(255, 255, 255, 0.18);
}

.home-icon::before {
  content: "⌂";
}

.help-icon::before {
  content: "?";
}

.nav-progress {
  display: flex;
  justify-content: flex-end;
  align-items: center;
  gap: 14px;
  color: var(--ink);
  font-weight: 700;
}

.nav-progress i {
  display: block;
  width: min(160px, 16vw);
  height: 9px;
  border-radius: 999px;
  background: linear-gradient(90deg, var(--accent-2) 0 58%, #dfe2db 58% 100%);
}

main {
  width: min(1480px, calc(100% - 32px));
  margin: 0 auto;
  padding: 10px 0 28px;
}

.hero {
  display: grid;
  grid-template-columns: minmax(250px, 0.58fr) minmax(620px, 1.72fr);
  grid-template-areas:
    "intro info"
    "intro key";
  gap: 10px;
  align-items: stretch;
}

.hero-copy,
.panel,
.quick-key,
.species-card,
.view-info-panel {
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: var(--radius);
  box-shadow: var(--shadow);
}

.hero-copy {
  grid-area: intro;
  align-self: stretch;
  display: flex;
  flex-direction: column;
  justify-content: flex-start;
  height: 100%;
  min-height: 100%;
  padding: clamp(12px, 1.8vw, 18px);
}

.hero-copy h2,
.section-title h2 {
  margin: 0;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(1.45rem, 3vw, 2.45rem);
  line-height: 1.02;
  letter-spacing: 0;
}

.hero-copy p:not(.eyebrow) {
  max-width: 58ch;
  color: var(--muted);
  font-size: 0.94rem;
  margin: 6px 0 0;
}

.intro-neuropter {
  margin: 9px 0 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--soft);
}

.intro-list {
  margin-top: 8px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: rgba(238, 241, 233, 0.72);
  padding: 8px 9px;
}

.intro-list h3 {
  margin: 0 0 4px;
  color: var(--accent-2);
  font-size: 0.82rem;
  text-transform: uppercase;
}

.intro-list ul {
  display: grid;
  gap: 3px;
  margin: 0;
  padding-left: 18px;
  color: var(--muted);
  font-size: 0.86rem;
}

.intro-list li::marker {
  color: var(--accent-3);
}

.intro-neuropter img {
  display: block;
  width: 100%;
  max-height: 132px;
  object-fit: cover;
}

.intro-neuropter figcaption {
  padding: 7px 9px;
  color: var(--muted);
  font-size: 0.72rem;
  line-height: 1.35;
}

.side-note {
  display: grid;
  grid-template-columns: 24px 1fr;
  gap: 10px;
  margin-top: 8px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: linear-gradient(135deg, var(--soft), rgba(248, 246, 241, 0.82));
  padding: 9px;
}

.side-note p {
  margin: 0;
}

.secondary-action {
  display: inline-flex;
  justify-content: center;
  align-items: center;
  width: fit-content;
  min-height: 38px;
  margin-top: 10px;
  border: 1px solid var(--line);
  border-radius: 999px;
  background: var(--cream);
  color: var(--accent-2);
  padding: 8px 14px;
  font-weight: 800;
  text-decoration: none;
}

.hero-copy .secondary-action {
  margin-top: auto;
}

.view-info-panel {
  grid-area: info;
  display: grid;
  grid-template-columns: 1.05fr 1.05fr 0.9fr;
  align-content: start;
  gap: 8px;
  padding: clamp(10px, 1.5vw, 14px);
}

.view-info-panel section {
  min-height: 100%;
  padding: 0 8px 0 0;
  border-right: 1px solid var(--line);
  border-bottom: 0;
}

.view-info-panel section:last-child {
  padding-right: 0;
  border-right: 0;
}

.view-info-panel h3 {
  display: flex;
  align-items: center;
  gap: 9px;
  margin: 0 0 6px;
  color: var(--accent-2);
  font-size: 0.82rem;
  text-transform: uppercase;
}

.view-info-panel p,
.view-info-panel ul {
  margin: 0;
  color: var(--muted);
}

.view-info-panel ul {
  display: grid;
  gap: 4px;
  padding-left: 20px;
}

.view-info-panel li::marker {
  color: var(--accent-3);
}

.learning-card {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: linear-gradient(135deg, var(--soft), var(--cream));
  padding: 10px;
}

.eyebrow {
  margin: 0 0 8px;
  color: var(--accent-2);
  font-size: 0.78rem;
  font-weight: 800;
  letter-spacing: 0;
  text-transform: uppercase;
}

.quick-key {
  grid-area: key;
  position: relative;
  z-index: 10;
  display: flex;
  flex-direction: column;
  padding: clamp(12px, 1.8vw, 18px);
  transition: transform 180ms ease, box-shadow 180ms ease;
  overflow: visible;
}

.photo-type-control {
  margin-top: 10px;
  color: var(--accent-2);
  font-size: 0.78rem;
  text-transform: uppercase;
  position: relative;
  z-index: 30;
  overflow: visible;
}

.native-photo-select {
  position: absolute;
  inline-size: 1px !important;
  block-size: 1px !important;
  width: 1px !important;
  min-height: 1px !important;
  border: 0 !important;
  padding: 0 !important;
  opacity: 0;
  pointer-events: none;
}

.custom-photo-select {
  position: relative;
  z-index: 40;
  display: block;
  margin-top: 6px;
  text-transform: none;
}

.custom-select-button {
  position: relative;
  z-index: 42;
  display: flex;
  justify-content: space-between;
  align-items: center;
  width: 100%;
  min-height: 50px;
  border-radius: var(--radius);
  background: var(--panel);
  color: var(--ink);
  font-weight: 800;
  text-align: left;
}

.custom-select-button::after {
  content: "⌄";
  color: var(--accent-2);
  font-size: 1rem;
}

.custom-select-options {
  position: absolute;
  z-index: 80;
  top: calc(100% + 4px);
  left: 0;
  right: 0;
  display: none;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
  box-shadow: 0 16px 36px rgba(34, 38, 35, 0.18);
}

.custom-photo-select.is-open .custom-select-options {
  display: grid;
}

.custom-select-options button {
  width: 100%;
  min-height: 38px;
  border: 0;
  border-bottom: 1px solid var(--line);
  border-radius: 0;
  background: var(--panel);
  text-align: left;
  font-weight: 800;
}

.custom-select-options button:last-child {
  border-bottom: 0;
}

.custom-select-options button[aria-selected="true"],
.custom-select-options button:hover {
  background: var(--soft);
  color: var(--accent-2);
}

.morph-image-frame {
  position: relative;
  overflow: hidden;
  background: #f4f1e7;
}

.morph-image-frame img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  transform: scale(1);
  transform-origin: 50% 45%;
  transition: transform 260ms ease, transform-origin 260ms ease;
}

.morph-image-frame.view-forewing img {
  transform: scale(3.35);
  transform-origin: 78% 18%;
}

.morph-image-frame.view-hindwing img {
  transform: scale(3.25);
  transform-origin: 79% 54%;
}

.morph-image-frame.view-head img {
  transform: scale(4.4);
  transform-origin: 16% 20%;
}

.morph-labels {
  position: absolute;
  inset: 0;
  pointer-events: none;
  opacity: 0;
  transition: opacity 180ms ease;
}

.learning-on .morph-labels {
  opacity: 1;
}

.morph-labels span {
  position: absolute;
  z-index: 3;
  padding: 4px 8px;
  border: 1px solid rgba(40, 89, 41, 0.25);
  border-radius: 999px;
  background: rgba(255, 252, 241, 0.92);
  color: var(--accent);
  font-size: 0.76rem;
  font-weight: 800;
  box-shadow: 0 8px 22px rgba(34, 38, 35, 0.12);
  transform: translate(-50%, -50%);
  white-space: nowrap;
}

.observe-box {
  padding: 10px 12px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--soft);
}

.observe-box h4 {
  margin: 0 0 8px;
  color: var(--accent);
  font-size: 0.82rem;
  text-transform: uppercase;
}

.observe-box ul {
  display: grid;
  grid-template-columns: repeat(4, minmax(140px, 1fr));
  gap: 6px 18px;
  margin: 0;
  padding-left: 18px;
  color: var(--muted);
}

.key-head,
.section-title {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 14px;
}

.key-head span,
.badge {
  display: inline-flex;
  align-items: center;
  min-height: 28px;
  border-radius: 999px;
  background: var(--soft);
  color: var(--accent-2);
  padding: 4px 10px;
  font-size: 0.82rem;
  font-weight: 800;
}

.progress-wrap {
  display: grid;
  gap: 8px;
  margin-top: 10px;
  color: var(--muted);
  font-size: 0.86rem;
  font-weight: 800;
}

.progress-track {
  height: 9px;
  overflow: hidden;
  border-radius: 999px;
  background: #deded4;
}

.progress-bar {
  width: 0;
  height: 100%;
  border-radius: inherit;
  background: linear-gradient(90deg, var(--accent-2), var(--accent-3));
  transition: width 220ms ease;
}

.quick-key h3 {
  margin: 12px 0;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(1.35rem, 3.1vw, 2.2rem);
  line-height: 1.18;
}

.key-visual {
  position: relative;
  height: 220px;
  margin-top: 12px;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: #f4f1e7;
}

.key-visual-image {
  position: absolute;
  inset: 0;
  background-image: url("assets/images/esquema-morfologic-libelloides.png");
  background-repeat: no-repeat;
  background-size: 104%;
  background-position: 50% 49%;
  transition: background-position 220ms ease, background-size 220ms ease;
}

.key-visual.view-forewing .key-visual-image {
  background-size: 178%;
  background-position: 50% 30%;
}

.key-visual.view-hindwing .key-visual-image {
  background-size: 178%;
  background-position: 51% 59%;
}

.visual-title {
  position: absolute;
  z-index: 2;
  font-size: 0.78rem;
  font-weight: 800;
  left: 12px;
  bottom: 12px;
  max-width: calc(100% - 24px);
  padding: 5px 9px;
  border: 1px solid rgba(47, 94, 59, 0.18);
  border-radius: 999px;
  background: rgba(255, 252, 241, 0.9);
  color: var(--accent);
  text-transform: uppercase;
  box-shadow: 0 8px 18px rgba(34, 38, 35, 0.1);
}

.answer-grid {
  display: grid;
  gap: 8px;
}

.answer-grid button {
  width: 100%;
  min-height: 40px;
  text-align: left;
  color: var(--ink);
  background: #fffdf8;
  transition: border-color 160ms ease, transform 160ms ease, background 160ms ease;
}

.answer-grid button:hover {
  border-color: rgba(47, 94, 59, 0.34);
  background: var(--soft);
  transform: translateY(-1px);
}

.key-controls {
  display: flex;
  flex-wrap: wrap;
  gap: 10px;
  margin-top: 12px;
}

.key-controls button {
  min-height: 40px;
  border-radius: 999px;
  font-weight: 800;
}

.key-controls button:disabled {
  cursor: not-allowed;
  opacity: 0.45;
}

.key-compatible {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(0, 1fr);
  gap: 10px;
  margin-top: 10px;
}

.key-compatible > div,
.key-compatible details,
.key-trail {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--cream);
  padding: 10px;
}

.key-compatible h4,
.key-trail h4 {
  margin: 0 0 8px;
  color: var(--accent-2);
  font-size: 0.88rem;
  text-transform: uppercase;
}

.key-compatible ul {
  display: grid;
  gap: 5px;
  margin: 0;
  padding-left: 18px;
  color: var(--muted);
}

.key-compatible li span {
  display: block;
  margin-top: 2px;
  font-size: 0.78rem;
}

.key-compatible summary {
  cursor: pointer;
  color: var(--accent-2);
  font-weight: 900;
}

.key-trail {
  display: grid;
  gap: 8px;
  margin-top: 12px;
  max-height: 220px;
  overflow: auto;
}

.key-trail p {
  margin: 0;
  border-top: 1px solid var(--line);
  padding-top: 8px;
  color: var(--muted);
}

.key-trail p:first-of-type {
  border-top: 0;
  padding-top: 0;
}

.key-trail b,
.key-trail span,
.key-trail small {
  display: block;
}

.key-trail span {
  color: var(--ink);
  font-weight: 800;
}

.panel {
  margin-top: 10px;
  padding: clamp(12px, 2.4vw, 18px);
}

.section-note {
  max-width: 72ch;
  margin: 8px 0 0;
  color: var(--muted);
}

.inaturalist-panel {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(280px, 0.65fr);
  gap: 14px;
  align-items: stretch;
  background:
    linear-gradient(135deg, rgba(248, 246, 241, 0.96), rgba(238, 241, 233, 0.92)),
    var(--panel);
}

.participation-copy {
  display: grid;
  align-content: start;
  gap: 8px;
}

.participation-copy h2 {
  margin: 0;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(1.65rem, 3vw, 2.55rem);
  line-height: 1.02;
}

.participation-copy p {
  max-width: 82ch;
  margin: 0;
  color: var(--muted);
}

.participation-lead {
  color: var(--ink) !important;
  font-size: 1.08rem;
  font-weight: 900;
}

.participation-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 10px;
  margin-top: 8px;
}

.participation-button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 8px;
  min-height: 44px;
  border: 1px solid rgba(47, 94, 59, 0.28);
  border-radius: 999px;
  background: rgba(255, 253, 248, 0.86);
  color: var(--accent-2);
  padding: 9px 15px;
  font-weight: 900;
  text-decoration: none;
  box-shadow: 0 8px 20px rgba(34, 38, 35, 0.07);
  transition: transform 160ms ease, border-color 160ms ease, background 160ms ease;
}

.participation-button.primary {
  background: var(--accent-2);
  border-color: var(--accent-2);
  color: #fffdf8;
}

.participation-button:hover {
  transform: translateY(-1px);
  border-color: rgba(47, 94, 59, 0.48);
  background: var(--soft);
}

.participation-button.primary:hover {
  background: #284f32;
  color: #fffdf8;
}

.participation-reasons {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: rgba(255, 253, 248, 0.72);
  padding: 12px;
}

.participation-reasons h3 {
  margin: 0 0 9px;
  color: var(--accent);
  font-size: 0.9rem;
  text-transform: uppercase;
}

.participation-reasons ul {
  display: grid;
  gap: 8px;
  margin: 0;
  padding: 0;
  list-style: none;
}

.participation-reasons li {
  display: grid;
  grid-template-columns: 26px 1fr;
  gap: 8px;
  align-items: center;
  border: 1px solid rgba(47, 94, 59, 0.1);
  border-radius: 10px;
  background: rgba(238, 241, 233, 0.7);
  padding: 8px 10px;
  color: var(--muted);
  font-weight: 700;
}

.id-form {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 12px;
  margin-top: 18px;
}

label {
  display: grid;
  gap: 6px;
  color: var(--muted);
  font-size: 0.9rem;
  font-weight: 700;
}

select {
  width: 100%;
  min-height: 50px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
  color: var(--ink);
  padding: 10px 14px;
  font: inherit;
  font-weight: 800;
  position: relative;
  z-index: 31;
  appearance: auto;
}

.result {
  margin-top: 16px;
  border: 1px solid var(--line);
  border-left: 5px solid var(--accent-3);
  border-radius: var(--radius);
  background: linear-gradient(135deg, var(--soft), var(--cream));
  padding: 16px;
}

.result.hidden {
  display: none;
}

.result strong {
  display: block;
  margin-bottom: 4px;
}

.result small {
  display: block;
  margin: 4px 0 10px;
  color: var(--muted);
  font-size: 0.84rem;
}

.table-wrap {
  overflow-x: auto;
  margin-top: 18px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  box-shadow: 0 12px 34px rgba(41, 45, 37, 0.05);
}

table {
  width: 100%;
  min-width: 980px;
  border-collapse: collapse;
  background: var(--panel);
}

th,
td {
  border-bottom: 1px solid var(--line);
  padding: 12px;
  text-align: left;
  vertical-align: top;
}

th {
  position: sticky;
  top: 0;
  background: #eef1e9;
  color: var(--accent-2);
  font-size: 0.86rem;
  text-transform: uppercase;
}

tr:last-child td {
  border-bottom: 0;
}

.species-grid {
  display: grid;
  grid-template-columns: 1fr;
  gap: 14px;
  margin-top: 12px;
}

.species-card {
  display: grid;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 16px;
  background: var(--panel);
  box-shadow: 0 14px 34px rgba(37, 43, 37, 0.08);
}

.species-header {
  display: grid;
  grid-template-columns: auto 1fr;
  gap: 14px;
  align-items: start;
  padding: clamp(12px, 2vw, 18px) clamp(12px, 2vw, 20px) 10px;
  border-bottom: 1px solid var(--line);
  background:
    linear-gradient(135deg, rgba(238, 241, 233, 0.86), rgba(255, 253, 248, 0.8)),
    var(--panel);
}

.species-status {
  display: inline-flex;
  align-items: center;
  gap: 8px;
  margin: 0;
  border: 1px solid rgba(47, 94, 59, 0.18);
  border-radius: 999px;
  background: var(--soft);
  color: var(--accent-2);
  padding: 5px 9px;
  font-size: 0.78rem;
  font-weight: 900;
  text-transform: uppercase;
  white-space: nowrap;
}

.species-header h3 {
  margin: 0;
  font-family: ui-serif, Georgia, "Times New Roman", serif;
  font-size: clamp(1.65rem, 3.4vw, 2.85rem);
  line-height: 1;
}

.species-authority {
  margin: 4px 0 0;
  color: var(--muted);
  font-size: 1.05rem;
}

.species-common-name {
  margin: 2px 0 0;
  color: var(--accent-2);
  font-weight: 800;
}

.species-layout {
  display: grid;
  grid-template-columns: minmax(360px, 46%) minmax(420px, 54%);
  gap: 0;
  align-items: start;
}

.species-media-panel {
  display: grid;
  align-content: start;
  align-self: start;
  padding: clamp(10px, 1.7vw, 16px);
  border-right: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
  background:
    linear-gradient(180deg, rgba(248, 246, 241, 0.9), rgba(238, 241, 233, 0.72));
}

.species-photo {
  display: grid;
  grid-template-rows: clamp(330px, 39vh, 470px) auto;
  height: auto;
  align-self: start;
  margin: 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 14px;
  background: var(--soft);
  box-shadow: 0 16px 42px rgba(38, 45, 35, 0.1);
}

.species-photo img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  filter: saturate(0.9) contrast(0.96);
  transition: transform 220ms ease;
}

.species-photo-button {
  position: relative;
  display: block;
  width: 100%;
  height: 100%;
  padding: 0;
  border: 0;
  background: transparent;
  cursor: zoom-in;
}

.species-photo-button:focus-visible {
  outline: 3px solid var(--accent-2);
  outline-offset: -3px;
}

.zoom-cue {
  position: absolute;
  right: 14px;
  bottom: 14px;
  border-radius: 999px;
  background: rgba(47, 94, 59, 0.9);
  color: #fff;
  padding: 7px 10px;
  font-size: 0.78rem;
  font-weight: 800;
}

.species-card:hover .species-photo img {
  transform: scale(1.025);
}

.species-photo figcaption {
  padding: 11px 14px;
  color: var(--muted);
  font-size: 0.82rem;
  background: rgba(255, 253, 248, 0.95);
}

.species-guide-panel {
  display: flex;
  flex-direction: column;
  padding: clamp(10px, 1.7vw, 16px);
}

.species-summary {
  margin: 0 0 9px;
  border-left: 5px solid var(--accent);
  border-radius: 0 14px 14px 0;
  background: linear-gradient(135deg, var(--cream), rgba(238, 241, 233, 0.72));
  color: var(--ink);
  padding: 10px 12px;
  font-size: clamp(0.95rem, 1.05vw, 1.02rem);
  font-weight: 650;
}

.species-info-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 8px;
  align-items: start;
}

.species-info-block {
  min-height: 0;
  border: 1px solid var(--line);
  border-radius: 10px;
  background: #fffdf8;
  padding: 10px;
  box-shadow: 0 10px 28px rgba(47, 55, 43, 0.045);
}

.diagnosis-block,
.observation-block {
  grid-column: 1 / -1;
}

.species-info-block h4 {
  display: flex;
  align-items: center;
  gap: 9px;
  margin: 0 0 7px;
  color: var(--accent-2);
  font-size: 0.88rem;
  font-weight: 900;
  text-transform: uppercase;
}

.species-info-block p,
.species-info-block ul {
  margin: 0;
  color: var(--muted);
}

.species-info-block ul {
  display: grid;
  gap: 4px;
  padding-left: 20px;
}

.species-info-block li::marker {
  color: var(--accent-2);
}

.warning-block li::marker {
  color: #b7831e;
}

.bibliography-block {
  background: linear-gradient(135deg, var(--soft), var(--cream));
}

.lightbox {
  position: fixed;
  inset: 0;
  z-index: 50;
  display: grid;
  place-items: center;
  padding: 22px;
  background: rgba(24, 28, 24, 0.72);
}

.lightbox.hidden {
  display: none;
}

.lightbox button {
  position: fixed;
  top: 18px;
  right: 18px;
  z-index: 2;
  border: 1px solid rgba(255, 255, 255, 0.35);
  background: rgba(255, 252, 241, 0.94);
  color: var(--ink);
}

.lightbox figure {
  width: min(1100px, 96vw);
  max-height: 92vh;
  margin: 0;
  overflow: hidden;
  border-radius: var(--radius);
  background: var(--panel);
  box-shadow: var(--shadow);
}

.lightbox img {
  display: block;
  width: 100%;
  max-height: 82vh;
  object-fit: contain;
  background: #111;
}

.lightbox figcaption {
  padding: 10px 14px;
  color: var(--muted);
  font-size: 0.86rem;
}

.facts {
  display: grid;
  gap: 8px;
  margin-top: 12px;
}

.fact {
  border-top: 1px solid var(--line);
  padding-top: 8px;
}

.fact b {
  display: block;
  font-size: 0.82rem;
}

.warning {
  background: var(--warn);
  border-radius: var(--radius);
  padding: 10px;
  color: #6a4a10;
}

.result-list {
  display: grid;
  gap: 16px;
  margin-top: 14px;
}

.result-species {
  display: grid;
  grid-template-columns: 1fr;
  gap: 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
}

.result-photo {
  display: grid;
  grid-template-rows: minmax(320px, 46vh) auto;
  margin: 0;
  overflow: hidden;
  border-bottom: 1px solid var(--line);
  background: var(--soft);
}

.result-photo img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  filter: saturate(0.9);
}

.result-photo figcaption {
  padding: 8px 14px;
  color: var(--muted);
  font-size: 0.78rem;
}

.result-species-body {
  padding: 18px;
}

.result-species h4 {
  margin: 0;
  font-size: clamp(1.8rem, 4vw, 3rem);
  line-height: 1;
}

.result-facts {
  display: grid;
  gap: 10px;
  margin-top: 14px;
}

.result-facts p,
.map-note {
  display: grid;
  grid-template-columns: 24px 110px minmax(0, 1fr);
  gap: 8px;
  align-items: start;
  margin: 0;
  color: var(--muted);
}

.result-facts b,
.map-note b {
  color: var(--ink);
}

.map-note {
  margin-top: 12px;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--soft);
  padding: 12px;
}

.info-icon {
  display: inline-block;
  width: 18px;
  height: 18px;
  margin-top: 3px;
  border: 2px solid var(--accent-2);
  border-radius: 50%;
  opacity: 0.82;
}

.info-icon.distribution {
  border-radius: 50% 50% 50% 0;
  transform: rotate(-45deg);
}

.info-icon.photo {
  border-radius: 4px;
}

.info-icon.diagnosis {
  border-radius: 2px;
  transform: rotate(45deg);
}

.info-icon.observation {
  border-radius: 50%;
  background: radial-gradient(circle at center, var(--accent-2) 0 3px, transparent 4px);
}

.info-icon.map {
  border-radius: 3px;
  border-color: var(--accent);
}

.info-icon.habitat {
  border-radius: 50% 50% 20% 20%;
  border-color: var(--accent-3);
  background: linear-gradient(135deg, transparent 45%, rgba(108, 155, 83, 0.25) 46%);
}

.info-icon.phenology {
  border-color: var(--accent);
  border-radius: 50%;
  background: conic-gradient(from 20deg, rgba(180, 138, 60, 0.3), transparent 42%, rgba(180, 138, 60, 0.24));
}

.reference-viewer {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(0, 1fr);
  gap: 12px;
  margin-top: 10px;
  align-items: start;
}

.photo-drop,
.photo-preview,
.reference-photo,
.reference-copy {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
}

.reference-photo {
  display: grid;
  grid-template-rows: clamp(330px, 31vw, 470px) auto;
  margin: 0;
  overflow: hidden;
}

.lateral-reference img {
  object-fit: cover;
  object-position: center;
}

.reference-photo img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  filter: saturate(0.86) contrast(0.96);
}

.reference-photo figcaption {
  padding: 8px 10px;
  color: var(--muted);
  font-size: 0.78rem;
}

.reference-copy {
  grid-column: 1 / -1;
  display: grid;
  grid-template-columns: minmax(260px, 0.8fr) minmax(420px, 1.35fr) minmax(220px, 0.75fr);
  align-items: stretch;
  gap: 9px;
  padding: 12px;
}

.reference-copy h3 {
  margin: 0;
  font-size: 1.2rem;
}

.reference-copy p {
  margin: 0;
  color: var(--muted);
}

.reference-copy > h3,
.reference-copy > p {
  grid-column: 1;
}

.reference-copy .observe-box {
  grid-column: 2;
  grid-row: 1 / span 3;
}

.reference-copy .photo-drop {
  grid-column: 3;
  grid-row: 1 / span 2;
}

.reference-copy .inat-link {
  grid-column: 3;
  grid-row: 3;
}

.photo-guide {
  margin: 12px 0 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: #fffdf8;
  box-shadow: var(--shadow);
}

.photo-guide img {
  display: block;
  width: 100%;
  height: auto;
  background: #fffdf8;
}

.photo-guide figcaption {
  padding: 8px 12px;
  border-top: 1px solid var(--line);
  color: var(--muted);
  font-size: 0.78rem;
}

.photo-drop {
  display: grid;
  place-items: center;
  align-content: center;
  min-height: 86px;
  padding: 12px;
  color: var(--ink);
  text-align: center;
  cursor: pointer;
}

.photo-drop input {
  position: absolute;
  inline-size: 1px;
  block-size: 1px;
  opacity: 0;
}

.photo-drop span {
  font-weight: 800;
}

.photo-drop small {
  max-width: 28ch;
  color: var(--muted);
}

.photo-preview {
  display: grid;
  place-items: center;
  grid-column: 1 / -1;
  min-height: 220px;
  overflow: hidden;
  color: var(--muted);
  background: var(--soft);
}

.photo-preview.hidden {
  display: none;
}

.photo-preview img {
  width: 100%;
  height: 100%;
  max-height: 420px;
  object-fit: contain;
}

.result.compact {
  margin-top: 0;
}

.inat-link {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 38px;
  border: 1px solid var(--accent-2);
  border-radius: var(--radius);
  color: var(--accent-2);
  font-weight: 800;
  text-decoration: none;
}

.morphology-layout {
  display: grid;
  grid-template-columns: 1fr;
  gap: 12px;
  margin-top: 10px;
  align-items: start;
}

.diagram-card {
  display: grid;
  grid-template-rows: clamp(430px, 58vw, 760px) auto;
  min-height: 0;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: #f4f1e7;
  overflow: hidden;
  margin: 0;
}

.diagram-card figcaption {
  padding: 9px 12px;
  border-top: 1px solid var(--line);
  color: var(--muted);
  font-size: 0.8rem;
  background: rgba(255, 252, 241, 0.86);
}

.morphology-grid {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(280px, 1fr));
  gap: 10px;
}

.morphology-section {
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
  padding: 10px;
}

.morphology-section h3 {
  margin: 0 0 6px;
}

.morphology-item {
  border-top: 1px solid var(--line);
  padding: 7px 0 0;
  margin-top: 7px;
}

.morphology-item div {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 8px;
}

.morphology-item span {
  border-radius: 999px;
  background: var(--soft);
  color: var(--accent-2);
  padding: 3px 8px;
  font-size: 0.74rem;
  font-weight: 800;
  white-space: nowrap;
}

.morphology-item p {
  margin: 6px 0 0;
  color: var(--muted);
}

.site-footer {
  display: flex;
  flex-wrap: wrap;
  justify-content: space-between;
  gap: 12px;
  border-top: 1px solid var(--line);
  padding: 22px clamp(16px, 4vw, 40px);
  color: var(--muted);
  font-size: 0.9rem;
  background: #ebe8dc;
}

.site-footer div {
  display: grid;
  gap: 4px;
  max-width: 620px;
}

.site-footer strong {
  color: var(--ink);
}

@media (max-width: 1180px) {
  .step-nav {
    grid-template-columns: repeat(4, auto);
    justify-content: start;
    overflow-x: auto;
  }

  .nav-progress,
  .help-pill {
    display: none;
  }

  .hero {
    grid-template-columns: minmax(230px, 0.58fr) minmax(480px, 1.42fr);
  }

  .view-info-panel {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .view-info-panel section {
    padding-bottom: 0;
    border-bottom: 0;
  }

  .reference-copy {
    grid-template-columns: minmax(220px, 0.75fr) minmax(360px, 1.25fr);
  }

  .reference-copy .photo-drop,
  .reference-copy .inat-link {
    grid-column: 1 / -1;
    grid-row: auto;
  }

  .reference-copy .photo-drop {
    min-height: 72px;
  }

  .inaturalist-panel {
    grid-template-columns: 1fr;
  }
}

@media (max-width: 900px) {
  .key-head,
  .section-title {
    align-items: flex-start;
    flex-direction: column;
  }

  .site-hero {
    min-height: 190px;
  }

  .hero-content {
    flex-direction: column;
  }

  .hero-brand {
    font-size: 1rem;
  }

  .hero-brand img {
    width: 132px;
  }

  .hero,
  .id-form,
  .reference-viewer,
  .result-species,
  .species-layout,
  .key-compatible,
  .morphology-layout,
  .species-grid {
    grid-template-columns: 1fr;
  }

  .hero {
    grid-template-areas:
      "intro"
      "info"
      "key";
  }

  .species-header {
    grid-template-columns: 1fr;
  }

  .species-media-panel {
    border-right: 0;
    border-bottom: 1px solid var(--line);
  }

  .species-photo {
    grid-template-rows: clamp(260px, 38vh, 360px) auto;
  }

  .hero {
    min-height: 0;
  }

  .view-info-panel {
    grid-template-columns: 1fr;
  }

  .view-info-panel section {
    padding: 0 0 10px;
    border-right: 0;
    border-bottom: 1px solid var(--line);
  }

  .view-info-panel section:last-child {
    padding-bottom: 0;
    border-bottom: 0;
  }

  .diagram-card {
    position: relative;
    top: auto;
  }

  .result-photo {
    grid-template-rows: minmax(240px, 42vh) auto;
  }

  .reference-photo {
    grid-template-rows: clamp(260px, 52vw, 390px) auto;
  }

  .reference-copy {
    grid-template-columns: 1fr;
  }

  .reference-copy > h3,
  .reference-copy > p,
  .reference-copy .observe-box,
  .reference-copy .photo-drop,
  .reference-copy .inat-link {
    grid-column: 1;
    grid-row: auto;
  }

  .observe-box ul {
    grid-template-columns: repeat(2, minmax(130px, 1fr));
  }

  .participation-actions {
    flex-direction: column;
  }

  .participation-button {
    width: 100%;
  }
}

@media (max-width: 520px) {
  main {
    width: min(100% - 18px, 1480px);
    padding-top: 12px;
  }

  .step-nav {
    padding: 10px;
  }

  .step-pill {
    min-height: 40px;
    padding: 7px 11px;
    font-size: 0.86rem;
  }

  .home-pill,
  .help-pill {
    display: none;
  }

  .hero-content {
    width: min(100% - 26px, 1480px);
  }

  .hero-content h1 {
    font-size: clamp(2.05rem, 12vw, 3.2rem);
  }

  .intro-neuropter img {
    max-height: 130px;
  }

  .hero-copy,
  .quick-key,
  .view-info-panel,
  .panel {
    padding: 11px;
  }

  .species-header,
  .species-media-panel,
  .species-guide-panel {
    padding: 11px;
  }

  .species-info-grid {
    grid-template-columns: 1fr;
  }

  .species-photo {
    grid-template-rows: clamp(220px, 32vh, 290px) auto;
  }

  .diagram-card {
    grid-template-rows: clamp(260px, 78vw, 430px) auto;
  }

  .observe-box ul {
    grid-template-columns: 1fr;
  }

  .key-visual {
    height: 190px;
  }

}